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Heat is one of the biggest threats to your transmission, especially when you are pushing your 2010–2024 Toyota 4Runner to its limits on the trail or towing a heavy load. Whether you are hauling gear for work, overlanding deep into the backcountry, or simply want peace of mind on daily drives, keeping transmission temperatures in check is critical. That's why we developed a direct-fit transmission cooler specifically for the fifth-generation 4Runner.
This cooler is designed to work alongside the stock in-tank transmission cooler, adding much-needed cooling capacity without compromise. The tube-and-fin core, combined with high-efficiency internal fins, ensures maximum heat transfer to keep transmission fluid temps under control. By reducing temperatures, this cooler helps extend the life of your transmission and reduces the risk of costly failures.
Every detail of this kit is built for convenience and reliability. It comes with everything needed for installation, including pre-cut hoses and fittings that connect seamlessly with OEM or Mishimoto components. Our transmission lines meet SAE J1532 standards, giving you confidence that every connection is strong, leak-free, and built to last. A robust powder-coated finish protects the cooler against the harshest environments, while precise engineering and testing in the USA ensure consistent performance.
Whether you are climbing rocky trails, pulling a trailer, or just want a more reliable daily driver, this transmission cooler delivers the durability and efficiency your 4Runner deserves. Like all our products, it comes backed by our Mishimoto Lifetime Warranty with Accident Protection, so you can focus on the journey ahead without worrying about your transmission.
Features and Benefits
Increased cooling capacity reduces stress when towing or off-roading
Designed to supplement and work in tandem with in-tank transmission cooler found in OEM and Mishimoto radiators for fifth-gen 4Runners
Tube-and-fin core with high-efficiency internal fins for superior heat transfer
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